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I have encountered this bug numerous times and tried to reproduce it. and it happens almost all of the time. even though im not 100% sure wich method it is.
While using the Lucius pattern Lasgun, when ur toughness is broken while ur in the animation of going into or of aim down sights u cant shoot the gun anymore. U have to switch weapons first and then u will be able to shoot again.
It also happened around 3-5 times while i was charging up the shot with a slower version where i couldt shoot anymore after toughness was broken while charging (wich cancels the charg and makes u stuck not beeing able to shoot)